Found in estuaries and saltmarshes around high-tide level either in brackish pools or on wet mud or sedges. Often found under stones or pieces of driftwood lying on marshy ground.

Sheltered places around high-tide usually out of water.Lives under stones and shingle, in rock crevices and sea caves and rarely among maritime plants and debris from saltmarsh vegetation.

Usually lives out of water, under stones and driftwood in muddy places at high-tide or in saltmarshes among seablite (Suaeda maritima) and sea purslane (Halimione portulacoides).

Lives in non-tidal lagoons and drainage ditches in slightly brackish water (2.5-8ppt).

Found in strongly brackish water (10-33ppt) in sheltered lagoons and marsh drains. Lives permanently submerged with species such as Enteromorpha, Zostera and Potamogeton.

Lives in low to moderate salinities (5-25ppt) in quiet estuaries, ponds behind shingle bars, and lagoons and drainage ditches in coastal marshes. Generally sheltered places without direct connection to the sea. Tends to live permanently submerged on mud or vegetation (especially Enteromorpha and Potamogeton).

Found in damp places such as damp woods, marshes, lake margins, flushes and dune slacks. Generally intolerant of human disturbance.

Frequently found in the roots of grass in dry calcareous places such as roadside banks, chalk/limestone hillsides, cliffs and screes and the hollows of stabilised dunes. Intolerant of shade and not usually found in woods.

Rather dry places. Most typically on or at the base of ivy-covered stone walls, usually shaded by trees but also recorded from rocks, ground litter in open woodland, hedge banks, natural cliffs and occasionally in fixed sand dunes.

Calcareous fens and marshes; often at the edges of lowland rivers and lakes. Normally lives close to the ground on sedges and grasses but may also be found on taller marsh vegetation such as Phragmites and Alnus. Not tolerant of human disturbance.