Janolus hyalinus (Alder & Hancock, 1854)

Author & date of last revision: Ian Smith on 5 August 2012
Antiopella hyalina; Antiopa hyalinus Alder & Hancock, 1854;

Description

Body (excluding appendages)

Up to 30 mm long. Dorsally brown  [2] with opaque white flecks which concentrate in places into patches of white, two of which may resemble false eyes anterior to rhinophores [1].

Cerata
Numerous. Arranged along sides and across anterior and posterior of body, completely surrounding, and sometimes partially concealing, the dorsum.
Many wart-like papillae on surface, and one on apex [11]. Translucent whitish, shading to hyaline green basally [2], hence J. hyalinus from Latin hyalus = glass green. Lateral cerata have a brown internal digestive gland extending half  way up, or less [11]. Those across the posterior, and those anterior to the rhinophores, have no digestive gland. Many small flecks and streaks of brown, white and yellowish white pigment on surface.

Rhinophores  [5]
Truncated apex. Numerous elongate rib shape warts orietated obliquely along the rhinophore. Brown with much yellowish white freckling, especially on warts. Similarly coloured large wrinkled caruncle between the bases of the rhinophores [8].

Head
Partly covered with cerata. Seen from below; a small oral veil, translucent white freckled yellowish white, extends laterally into oral tentacles [3]. Mouth yellowish white. Small eye at base of each rhinophore.

Foot
Broad until posterior of body, from where the projecting metapodium tapers rapidly [4]. Bilaminate anterior edge smoothly rounded with no propodial lobes or tentacles. Sole transparent whitish, clearly revealing internal anatomy and the white freckles on the dorsal surface of the foot [3]

Key identification features

  • Cerata continue round anterior of head [1].
  • Caruncle between bases of rhinophores [8].
  • Many wart-like papillae on surface of cerata [11].

Similar species

Janolus cristata

  • Surface of cerata smooth

Proctonotus mucroniferus 

  •  No caruncle between rhinophores.

Ecology and behaviour

Most records sublittoral, on or near its erect bryozoans prey; species of  Bugula  (EML) and  Scrupocellaria (UWN). When contracted, resembles a sea anemone [6]. Both in extreme exposure (e.g. St Kilda) and extreme shelter (e.g. Lough Hyne). A relatively agile species with near neutral buoyancy, so very occasionally drifts on shore where it may be found in pools or in shallow water. Simultaneous hermaphrodite. Spawn a coiling line of white ovoids, each of which contains many ova [9], recorded September in wild, and May in refrigerator at 9° C.

Distribution and status

Mediterranean to northern Scotland (GBIF map ). One of the rarest British nudibranchs (Thompson & Brown, 1984). Scattered records from northern Scotland to English Channel, but not recorded in North Sea. ( UK interactive map )

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Lateral view of Janolus hyalinus. Length 15mm. Foot nearly transparent with pigment flecks on dorsal surface. Metapodium extending from posterior tapers rapidly to fine point. In water’s edge, LWS, Menai Strait, Wales. April 2011.
Image © I.F. Smith
Janolus hyalinus. Length 15mm. Brown dorsum marked with opaque white flecks which concentrate in places into patches of white; two  in front of rhinophores resemble false eyes. Floating in water’s edge, LWS, Menai Strait. April 2011.
Image © I.F. Smith

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Recorded UK distribution